Tomarus rostratus Dupuis 2014
- Dataset
- Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision
- Rank
- SPECIES
Classification
- kingdom
- Animalia
- phylum
- Arthropoda
- class
- Insecta
- order
- Coleoptera
- family
- Dynastidae
- genus
- Tomarus
- species
- Tomarus rostratus
description
(Figs. 5 F, 7 H, 9 L, 13 J, 14 F, 16 P, 20 G, 22 B, 28 G; 49)
description
Description. Habitus as in Fig. 28 G. Length 17.4 – 27.5 mm; humeral width 8.3 – 10.0 mm. Color dark reddish brown. Head: Frons coarsely and irregularly rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus trapezoidal, not strongly narrowed toward apex; apex with 2 triangular teeth separated by about 1.5 tooth diameters. Mandible with 2 apical teeth and a lateral, widely rounded tooth (Fig. 5 F). Mentum abruptly constricted at apical 3 rd. Galea of maxilla with teeth 5 and 6 well developed. Pronotum: Surface punctate, punctures rounded, smaller than punctures on elytra; densely distributed on apical and lateral margins. Apical margin with a small tubercle, visible in lateral view. Pronotum without rugose fovea, with small to large punctures on each side of tubercle. Scutellum: Surface without punctures. Elytra: Sutural stria complete. Elytral punctures dense and deep, punctures mainly ocellate, 3 pairs of distinct double rows. Abdomen: Pygidium convex (male) to flat (female) in lateral view; surface deeply rugopunctate on basal 3 rd with disc smooth (male) (Fig. 9 L), or completely rugopunctate (female). Apical margin of sternite VII straight in male, slightly emarginate with notch broad and shallow in female. Apex of sternite VIII emarginate, notch wide in male and narrow and deep in female. Venter: Prosternal process long, thick, apex transversely oval to rounded, and bordered with long, pale yellow setae. Metasternum with short, sparse setae on anterior angles. Legs: Protibia tridentate with an additional small, basal denticle. Apex of metatibia slightly crenulate, with 8 – 15 spinules. Female genitalia: Coxite surface strongly concave (Fig. 13 J). Male genitalia: Spiculum gastrale with basal plates (Fig. 14 F). Parameres widened at apical 4 th, elongate from basal 4 th to apex; sides parallel, without teeth or lateral projections (Fig. 16 P), regularly curved at lateral view (Fig. 20 G). Internal sac with small copulatory lamella, lamellar spiny belt reduced, complex of accessory lamellae with 6 spines joined at the base by dense granules (Fig. 22 B).
diagnosis
Diagnosis. Tomarus rostratus is unique in the genus because of its extremely elongate parameres, without lateral teeth. It can be also differentiated by the following combination of characters: clypeus trapezoidal; pronotum with small tubercle and without rugose fovea, with small to large punctures on each side of tubercle (Fig. 28 G); protibia tridentate with a small denticle; apical border of pygidium strongly truncate in male; apical margin of sternite VIII emarginated in female; coxite surface strongly concave (Fig. 13 J); spiculum gastrale with basal plates (Fig. 14 F); internal sac of aedeagus with small copulatory lamella and reduced lamellar spiny belt (Fig. 22 B).
distribution
Distribution. Tomarus rostratus is an endemic species from the coastal desert of Peru (Dupuis 2014) and Chile (López-García & Deloya 2018). Locality records (Fig. 49). 39 examined specimens from FSCA, MEKRB, MNHN, USNM, SENASA, UGBC. CHILE (12). Arica y Parinacota: Cuya; Puente Chapa (50 km S Arica); Valle de Azapa (6 km E Arica). PERU (27). Arequipa (3): Camaná, Quilca; Quiroz. Lima (19): Canta, Santa Rosa de Quives; Cañete; Huarochirí, Santa Eulalia; Huaura, Végueta, Medio Mundo; Lima. No data (5): Peru. Natural history. Specimens have been found between 12 – 1,036 m elevation between August and February.
materials_examined
Male holotype (MNHN) “ Peru. / Tomarus / Er. / Maimon // MUSÉUM PARIS / 1952 / coll. R. OBERTHÜR // Tomarus rostratus n. sp. / HOLOTYPE / F. Dupuis 2014 // HOLOTYPE // MNHN / EC 4187 ”. Type locality: Peru.
Name
- Homonyms
- Tomarus rostratus Dupuis 2014