Chassalia northiana T. Y. Yu 2021
- Dataset
- Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species
- Rank
- SPECIES
Classification
- kingdom
- Plantae
- phylum
- Tracheophyta
- class
- Magnoliopsida
- order
- Gentianales
- family
- Rubiaceae
- genus
- Chassalia
- species
- Chassalia northiana
description
Description Herb, shrub to small tree, 1 – 2 m tall, glabrous. Stem terete, young parts of stems slightly flattened and canaliculate when dry, internode 3 – 8 cm × 2 – 4 mm. Leaf-blades obovate to elliptic, usually broadest at ¼ to ½ length from apex, 12 – 27 × 4 – 9 cm, apex acuminate with acumen 1 – 1.5 (– 3) cm long, base truncate, midrib slightly raised above from both adaxial and abaxial, secondary nerves 8 – 14 on each side of the midrib, petioles 1 – 6 cm long. Stipule flabellate, margin usually broken, sometimes slightly bilobed or incised, 8 – 14 × 5 – 14 mm, with standard colleters inside. Inflorescence compound cyme, terminal, usually with rachis and 2 branches, 6 – 14 cm long in total. Peduncle 4 – 7.5 cm long, branches and first internode of rachis 1 – 4 cm long, branches usually 0.5 – 1 cm shorter than rachis, bracts of branches and first internode of rachis usually 4, separate or sheathed at base, 2 leaf-like bracts 1 – 1.8 cm × 1.5 – 2 mm, sometimes growing larger than young but not fully developed leaves 3 – 7 × 1.5 – 2 cm, 2 stipule-like bracts 5 – 8 × 5 – 8 mm, rachis and branches sometimes with second branches and second internode of rachis 2 – 10 mm long, capitular bracts 8 – 10 × 2 – 5 mm. Flowers sessile, 15 – 18 mm long, corolla tube lower part 0.6 – 1 mm wide, upper part 2 – 2.5 mm wide, mouth 3.5 – 4 mm wide, corolla lobes usually 5, 3 × 2.5 mm, stamens 2.3 – 2.5 mm long, inserted 1 mm from corolla mouth, inside corolla tube a band of hairs 0.7 mm long at base of stamens, hairs erect, colourless, 0.1 – 0.2 mm long; stigma bilobed, each lobe round, 0.5 mm in diameter; disc 1.2 × 1.2 mm. Hypanthium 4 × 2 mm, calyx tube 1 × 4 mm with 5 lobes 0.1 × 0.5 mm. Fruit black or deep brown when dry, blue when ripe, ellipsoid to round, 7 × 6 × 6 mm, calyx persistent, not accrescent, 0.5 × 2 mm, disc persistent, not accrescent, 1.2 × 1.2 mm. Pyrene plano-convex, 7 × 5.5 – 6 × 2 – 3 mm, dorsal surface convex, with two adjacent parallel longitudinal crests, 0.4 mm wide, 0.25 mm apart along their length, forming a longitudinal groove from apex to base between the two crests; ventral surface concave, central part slightly elevated, margin with an endocarpal outgrowth 1 mm wide, with a groove between the edge of endocarp and endocarpal outgrowth 0.5 mm wide; raphal opening at base of the endocarpal outgrowth, round, 0.4 mm wide, 0.2 – 0.3 mm from the edge of the pyrene wall, apex with a bony spine, spine triangular, erect, 0.8 × 0.2 mm. PGS not detected. Seed light brown, hemi-ellipsoid, 5 × 6 mm, longitudinally slightly crested at midline of dorsal surface, endosperm not detected.
description
urn: lsid: ipni. org: names: 77215414 - 1 Figs 28 – 29
diagnosis
Diagnosis Differs from Chassalia blumeana Govaerts to which it is most similar, by having inflorescence branches and rachis Ẑ 5 mm, usually 1 – 5 cm long, bracts of inflorescence branches and rachis usually consist of 2 leaf-like bracts and 2 stipule-like bracts, leaf-like bracts sometimes develop to 3 – 7 cm long.
discussion
Remarks This species is one of the most widespread of the Bornean ‘ involucrate’ species.
distribution
Distribution, habitat Borneo: Malaysia, from central Sabah to west Sarawak. Alluvial forest, stream banks, Kerangas forest. Lowland to montane forest, alt. 200 – 1000 m.
etymology
Etymology Named for Marianne North (1830 – 1890), who included this plant in painting 624 entitled Curious Plants from the Forest of Matang, Sarawak, Borneo (https: // images. kew. org / botanical-art / mariannenorth / 624 - curious-plants-forest-matang-sarawak- 5122480. html) which is the first image record of this species, made in the 1860 s. It is possibly the earliest record of the genus from Borneo.
materials_examined
Type MALAYSIA – Borneo, Sarawak • First Division, Kuching, Matang, proposed Matang National Park, Sungai Sendok; 8 May 1987; Bernard Lee Meng Hock S 53828; holotype: K [K 001129725]!; isotype: SAR. Additional material MALAYSIA – Borneo, Sabah • Ranau, Mininsalu; 22 Sep. 1988; Amin & Prancis SAN 123432; K, SAN. – Borneo, Sarawak • Sg. Rayu, Matang Forest Reserve; 27 Mar. 1988; Munting & Jugah, S 55620; K!, SAR • 5 th Division, along Sg. Maisa in Maligan Range, Lawas; alt. 3400 ft; 13 Mar. 1973; Ilias Paie S 32900; K!, SAR • Sri Aman Division, Temawai Berau, Sungai Selepong; 6 Apr. 1990; Rena, Runi, Rantai et al. S 58511; K!, SAR.
Name
- Homonyms
- Chassalia northiana T. Y. Yu 2021