Georissus Latreille, 1809

Dataset
GBIF Backbone Taxonomy: Rank
GENUS: Published in
Gen. Crust. Ins., 4

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Classification

kingdom
Animalia: phylum
Arthropoda: class
Insecta: order
Coleoptera: family
Georissidae: genus
Georissus

description

Supplementary description BODY. Black to reddish brown, oval, strongly convex dorsally. CLYPEUS. Uneven, with granules or carinae on its surface; articulated carinae arising at antero-lateral portion of clypeus and reaching anterior portion of frons (slr in Fig. 1 A); a pair of distinct bulges (smb in Fig. 1 A) on antero-median portion of frons (in Nipponogeorissus and Neogeorissus); sides of frons smooth, lacking granules (lsa in Fig. 1 A). Eyes developed, not protuberant. Antennae (Fig. 4 M – O) short, 9 - segmented, with 3 - segmented pubescent club (7 - segmented with one-segmented club in Australian species: Fikáček 2019). Mandibles (Fig. 4 J – L) large and well sclerotized, partly concealed beneath clypeus and labrum; antero-lateral margin forming obtuse angle; apex pointed; internal margin with microtrichia except apex. Maxillary palpi (Fig. 4 A – C) 4 - segmented; palpomere 4 longest and nearly same length as palpomeres 1 – 3 combined. Labrum (Fig. 4 D – F) semicircular, widest at base; lateral margins fringed with spines. Mentum subtriangular, strongly narrowed anteriorly, with long setae on disc. Labial palpi (Fig. 4 G – I) as long as mentum; apical palpomere oval to oblong oval, densely bearing spines in interior margin. PRONOTUM. Half oval to trapezoidal, divided into anterior and posterior portions; anterior portion irregularly arranged with granules or short grooves and pits, with shallow median longitudinal groove (smlg in Fig. 1 B); posterior portion more convex than anterior portion, with large fovea on anteromedian portion (cd in Fig. 1 B); posterior sides of cd with a pair of distinct foveae (sdp in Fig. 1 B) (only in Neogeorissus); sublateral potion of posterior portion with distinct bulge (slb in Fig. 1 B); behind of slb with deep groove (pg in Fig. 1 B); sides somewhat protruding laterally, having dentation (sdn in Fig. 1 B) in Neogeorissus. LEGS. Protrochantins concealed. Procoxae large, plate-like, concealing prosternum. Femora somewhat flattened. Tibiae slender, especially protibiae rather flattened. Tarsal formula 5 - 5 - 5 (5 - 4 - 4 in Australian species: Fikáček 2019). ELYTRA. Provided with ten punctate rows. All or alternate intervals slightly to highly raised. Hind wings well developed or absent in some species. VENTRITES. Ventrites 1 and 2 connate; ventrite 1 large, with distinct granules; posterior margin of ventrite 1 and anterior margin of ventrite 2 with a pair of ventral teeth (vt in Fig. 8 A). MALE GENITALIA (Fig. 9). Aedeagus trilobate type, almost symmetrical. FEMALE GENITALIA. Gonocoxite (Fig. 10) long, almost cross each other posteriorly. Gonostylus short, provided with two long setae at apex. Median sclerite unilobed, densely bearing long setae. Spermatheca (Fig. 11 E – H) slightly sclerotized, elongated oval. Bursal sclerite (Fig. 11 A – D) somewhat developed in subgenus Neogeorissus, slightly asymmetrical.

description

Redescription HEAD. Clypeus uneven, with groove along the anterior margin. Surface of clypeus provided with large granules densely; slr reaching to anterior margin of frons. Frons bearing median longitudinal groove to vertex; lsa narrow, not connecting each other. Granulation of frons sparse, indistinct. Mandibles (Fig. 4 J) relatively small; median lateral margin curving strongly, apex bidentate, mola well developed. Antennae (Fig. 4 M) relatively long; antennomere 4 as wide as antennomere 3, about twice as long as antennomere 3; pubescence of club short. Labrum (Fig. 4 D) not protruding anteriorly; spines on anterior margin indistinct. Apical labial palpomere (Fig. 4 G) oblong oval, with long spines densely. Galea (Fig. 4 A) bearing long dense setae. PRONOTUM. Half oval, slightly projecting laterally in lateral portion, without sdn. Lateral margin except anterior portion regularly provided with granules. First half of pronotal disc irregularly bearing foveae and grooves, the second half of disc smooth, but with median longitudinal groove reaching near posterior margin. E LYTRA. Bearing seriate large punctures. Elytral interstices smooth, not highly raised. Humeral bulges distinct, smooth. Lateral ridges on each elytron indistinct. ABDOMEN. Ventrites sparsely bearing granules. Ventrites 1 – 2 provided with a pair of ventral teeth. MALE GENITALIA. Aedeagus wide, subparallel-sided. Parameres densely provided with small spines on antero-internal portion, as long as phallobase. Median lobe little shorter than parameres. Basal struts short. Phallobase wide; basal foramen large, but indistinct. FEMALE GENITALIA. Ovipositor relatively elongate to longitudinal portion. Two long setae of gonostylus long.

discussion

Remarks This subgenus is distinguished by the following characteristics: lacking sdn (present in Neogeorissus); posterior portion of pronotal disc smooth, with median longitudinal groove (uneven, with granules or depression in other subgenera); elytral interstices smooth, not highly elevated (in contrast to uneven, granulated in other subgenera); parameres provided with small dense spines on the anterior portion (even in other subgenera). The members of this subgenus are often covered with sand or mud when they are alive (Bameul 1989; Shepard 2003; Litovkin 2018; Fikáček 2019). Bameul (1989) reported that adults of G. crenulatus attach sand grains to their body surface using oral secretions and their legs.

Name

Synonyms: Cathammistes Illiger, 1807; Georyssus Latreille, 1809; Georyssus Stephens, 1828; Neogeorissus Satô, 1972; Nipponogeorissus Satô, 1972
Homonyms: Georissus Latreille, 1809

Bibliographic References

Australian Faunal Directory, 2007 version. Available online at https://biodiversity.org.au/afd/home.
Australian Faunal Directory, 2007 version. Available online at https://biodiversity.org.au/afd/home.
Freude, et al. (1971)
Hallan, J. (2000-current). Biology Catalog (2012 version), http://bug.tamu.edu/research/collection/hallan/ (as at 2012).
Hansen, M. (1999) World Catalogue of Insects 2: Hydrophiloidea (Coleoptera). Apollo Books, Stenstrup, 416 pp.
International Commission on Zoological Nomenclature. Opinion 1891 Crenitis Bedel, 1881, Georissus Latreille, 1809 and Oosternum Sharp, 1882 (Insecta, Coleoptera): conserved.Bulletin of Zoological Nomenclature 55, 58-60 (1998)
Klausnitzer, B. (1994)
Latreille, P. A. (1809) Genera Crustaceorum et Insectorum, secundum ordinem naturalem in Familias disposita, iconibus exemplisque plurimis explicata. Vol. 4. 399 pp. - Amand Koe ni g, Paris iis et Argentorati. https://www.biodiversitylibrary.org/page/42114245
Latreille, P.A. (1809). Genera crustaceorum et insectorum. Tomus quartus et ultimus. Amand Koenig, Parisiis et Argentorati, 399 pp.
Mosquera Murillo, Z., Sánchez Vázquez, S.P. (2018) Riqueza genérica y distribución de los coleópteros acuáticos del Departamento del Chocó – Colombia, depositados en la Colección Limnológica de la Universidad Tecnológica del Chocó. Rev. U.D.C.A Act. & Div. Cient. 21(1): 155-165. https://doi.org/10.31910/rudca.v21.n1.2018.674
Murillo, Z. M., Mosquera, M. M. M. (2017) Diversidad de la entomofauna acuática y calidad de agua en quebradas del río San Juan, Chocó – Colombia. Revista U.D.C.A Actualidad & Divulgación Científica 20: 149–161. https://doi.org/10.31910/rudca.v20.n1.2017.72 https://doi.org/10.31910/rudca.v20.n1.2017.72
Neave, S. A. (1939-1996). Nomenclator Zoologicus vol. 1-10 Online. [developed by uBio, hosted online at MBLWHOI Library].
Neave, S. A. (1939-1996). Nomenclator Zoologicus vol. 1-10 Online. [developed by uBio, hosted online at MBLWHOI Library].
Nilsson, A.N. (1996)
as per family