Pterostichus Bonelli, 1810

Dataset
GBIF Backbone Taxonomy: Rank
GENUS: Published in
Bonelli, F.A. Observations entomologiques. Première partie (Cicindélètes et portion des carabiques) [including "Tabula Synoptica exhibens genera carabicorum in sectines et stirpes disposita"]. 58 pp. + 1 table. (Torino). (1810).

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Classification

kingdom
Animalia: phylum
Arthropoda: class
Insecta: order
Coleoptera: family
Carabidae: genus
Pterostichus

Pterostichus Bonelli 1810

Genus Pterostichus Bonelli, 1810

Pterostichus Bonelli, 1810: Tabula Synoptica. Type species: Carabus fasciatopunctatus Creutzer, 1799 designated by Curtis (1828: plate 196). Etymology. From the Greek pteron (wing, by extension elytron) and stichos (row, line), possibly alluding to the striae on the elytra of the adults [masculine].

Diversity.

About 1,055 species (Lorenz 2005: 269-287, 289, as Pterostichus, Lyropedius, and Abacidus) in the Nearctic (about 180 species, of which four are adventive) and Palaearctic (about 875 species) Regions. Percolaus Bates with two Middle American species, Mayaferonia Ball and Roughley with two Middle American species, and Allotriopus Bates with eight Mexican species are also includes in the genus Pterostichus by some authors (e.g., Ball and Roughley 1982).

Faunistic Note.

Pterostichus (Steropus) madidus (Fabricius), a European species, is known in North America from two specimens, one collected in 1913 in Quebec (Lindroth 1966: 476) and the other one in 1984 in Somerset County, Pennsylvania (Cameron and Reeves 1990: 127). Until more specimens are collected on this continent, the species is not considered as a North American resident.

Taxonomic Note.

Based on phylogenetic analyses from two nuclear gene sequences (wingless and 28SrDNA), Sasakawa and Kubota (2007) concluded that the genus Pte rostichus sensu Bousquet (1999) was monophyletic with the exception of Bothriopterus which forms a clade with Stomis and Myas.

Pterostichus Bonelli 1810

Genus Pterostichus Bonelli, 1810

Subgenus Argutor Dejean, 1821

Pterostichus (Argutor) cryobioides Chaudoir, 1868

Citations: Mountain area along the Iranian-Turkish border (Prov. of West Azarbaijan; Muilwijk, 2015).

Pterostichus (Argutor) cursor (Dejean, 1828)

Citations: Ziba-Kenar (Prov. of Gilan; Gueorguiev, 2011).

Pterostichus (Argutor) vernalis Panzer, 1796

Citations: Kurusch-göl (Prov. of East Azarbaijan; Mandl, 1963).

Subgenus Adelosia Stephens, 1835

Pterostichus (Adelosia) macer macer (Marsham, 1802) Citations: Gorgan env. (Prov. of Golestan; Afshari & Khormali, 2014).

Subgenus Melanius Bonelli, 1810

Pterostichus (Melanius) elongatus (Duftschmid, 1812)

Studied material: (ZMTU): NowShahr, 2010 (Prov. of Mazandaran) and Astaneh, 2012 (Prov. of Gilan). Citations: Ghaenat (Prov. of Razavi Khorasan; Sadeghi et al., 2010).

Pterostichus (Melanius) nigrita (Fabricius, 1792)

Citations: Kurusch-göl (Prov. of East Azarbaijan; Mandl, 1963).

Subgenus Bothriopterus Chaudoir, 1835

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787) = melanoscelis (Marseul, 1880)

Citations: Without exact locality (Bousquet, 2003 f).

Subgenus Platysma Bonelli, 1810

Pterostichus (Platysma) niger niger (Schaller, 1783)

= subcordatus Chaudoir, 1842

Studied material: Khirood Forest, 2012 (Prov. of Mazandaran); (ZMTU): Astaneh, 2012 (Prov. of Gilan). Citations: Without exact locality (Bousquet, 2003 f); Khirood forest, NowShahr (Prov. of Mazandaran; Azadbakhsh et al., 2015).

Subgenus Pseudomaseus Chaudoir, 1838

Pterostichus (Pseudomaseus) anthracinus anthracinus (Illiger, 1798) Citations: Without exact locality (Hieke & Wrase, 1988).

Pterostichus (Pseudomaseus) anthracinus depressiusculus Chaudoir, 1844 Citations: Province Mazandaran (Chaudoir, 1842).

Pterostichus (Pseudomaseus) fuscicornis (Reiche & Saulcy, 1855) Studied material: (ZMTU): Shemshak, 2008 (Prov. of Tehran), NowShahr, 2010 (Prov. of Mazandaran) and Namin, 2010 (Prov. of Ardabil).

Citations: Bludan NW of Damascus (Prov. of Lurestan; Angus et al., 2009); Without exact locality (Gueorguiev, 2011).

Subgenus Phonias Gozis, 1886

Pterostichus (Phonias) strenuus (Panzer, 1796)

Studied material: Khirood Forest, 2012 (Prov. of Mazandaran). Citations: Galhe rod-Khan Forest (Prov. of Gilan; Gougheri, 2014).

Subgenus Falsargutor Kryzhanovskij, 1983

Pterostichus (Falsargutor) pseudopedius iranianus Straneo, 1983 Studied material: (ZMTU): Nowshahr, 2008 (Prov. of Mazandaran). Citations: Without exact locality (Bousquet, 2003 f).

Subgenus Lyrothorax Chaudoir, 1838

Pterostichus (Lyrothorax) caspius (Ménétriés, 1832)

Studied material: Khirood Forest, 2012 (Prov. of Mazandaran). Citations: Basmenj Village (Prov. of East Azarbaijan; Atamehr, 2013).

Pterostichus

Key to subgenera of the genus Pterostichus from China (part)

1. Mesofemora with two or three setae on posterior margin; metaepisterna short, length subequal to the width of anterior margin; metacoxa with only two setae; hind trochanter without setae; penultimate labial palpomere without seta near apex; pronotum

with hind angles distinct, posterior seta placed near hind angle.................................................. 2 - Without the combination of above characters.......................................... Pterostichus other subgenera

2. Hind trochanter without seta............................................................................. 3

- Hind trochanter with one seta...................................................... Pterostichus other subgenera

3. Third interval of elytron with three or more setigerous pores, 5 th interval without pores; ridge between outer basal fovea of pronotum and lateral margin strongly carinate; right paramere thick.................. Pterostichus (Feroperis Lafer 1979)

- Third interval of elytron usually with two or less setigerous pores; if 3 rd interval has more than three pores, then 5 th interval has some pores; ridge between outer pronotal basal and lateral margin strongly or at most moderately convex; right paramere lamina............................................................................................. 4

4. Apical lamella of male genitalia laminar, rather wide and long, usually slightly twisted; species from the Himalayan Region....................................................................... Pterostichus (Pseudethira Sciaky, 1996)

- Male genitalia not like so; species not from the Himalayan Region............................................... 5

5. Umbilical series on 9 th interval continuous; apical lamella of male genitalia long and strongly twisted................................................................................... Pterostichus prattii Bates, 1890 species group

- Umbilical series on 9 th interval discontinuous; apical lamella of male genitalia at most slightly twisted.................. 6

6. Apical lamella of male genitalia wide, more or less bent dorsally; pronotal disc impunctate; females with strong isodiametric microsculpture on elytra; setae on terminal sternum distant from apex................. Pterostichus (Wraseiellus n. subg.)

- Apical lamella of male genitalia narrow and short, not bent dorsally; pronotum densely and finely punctate; setae on terminal sternum close to apex. (only applicable for Nialoe species from China)............... Pterostichus (Nialoe Tanaka, 1958)

Name

Synonyms: Aachnoideus Chaudoir, 1838; Abaxodes Gistel, 1857; Abea Morita, 1992; Adelopterus Reitter, 1886; Adelosia Stephens, 1835; Agonodemus Reitter, 1896; Agonodemus de Chaudoir, 1838; Agonoderus Gemminger & Harold, 1868; Alecto Heyden, 1883; Americomaseus Csiki, 1930; Amolops Tschitscherin, 1898; Amolopsa Strand, 1936; Anagodemus Motschoulsky, 1845; Anilloferonia Van Dyke, 1926; Anomostichus Sciaky, 1995; Aphaon Heyden, 1906; Arachnoidius de Chaudoir, 1838; Arguta Lucas, 1846; Argutor Stephens, 1827; Asioplatysma Kryzhanovskii, 1968
Homonyms: Pterostichus Bonelli, 1810; Pterostichus Broun; Pterostichus Stephens, 1827

Bibliographic References

Bonelli, F.A. (1810). Observations entomologiques. Première partie (cicindélètes et portion des carabiques) [with the "Tabula synoptica exhibens genera carabicorum in sectiones et stirpes disposita"]. Turin. 58 pp. + 1 table. [preprint of Mémoires de l'Académie Impériale des Sciences, Littérature et Beaux-Arts de Turin: Sciences Physiques et Mathématiques 1809-10(Mém. Prés.): 21-78, but the later version is missing the Tabula].
Bousquet, Y. (2012). Catalogue of Geadephaga (Coleoptera: Adephaga) of America, north of Mexico. <em>ZooKeys.</em> 245: 1-1722. 10.3897/zookeys.245.3416
Bousquet, Y. (2012). Catalogue of Geadephaga (Coleoptera: Adephaga) of America, north of Mexico. <em>ZooKeys.</em> 245: 1-1722. 10.3897/zookeys.245.3416
Bousquet, Yves, 2012: Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys, issue 245. 1-1722.
Bousquet, Yves, and André Larochelle, 1993: Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America North of Mexico. Memoirs of the Entomological Society of Canada, no. 167. 397.
CoL2006
Fauna Europaea (May 2010)
Hallan, J. (2000-current). Biology Catalog (2012 version), http://bug.tamu.edu/research/collection/hallan/ (as at 2012).
Lundberg & Gustafsson (1899-12-31 23:00:00) Catalogus Coleopterorum Sueciae
SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/.
SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/.
WoRMS (Mar 2013)
WoRMS (Mar 2013)
as per family (nonmarine except where stated)