Progonyleptoidellus Piza 1940
- Description of two new species of Progonyleptoidellus (Opiliones: Gonyleptidae), with a cladistic analysis of the genus, an overview of relationships in the K 92 group, and taxonomic notes on Deltaspidium
Diagnosis. Progonyleptoidellus can be distinguished from the other genera of Progonyleptoidellinae by the following combination of characters: Dorsal scutum outline (Figs. 2 – 7) gamma pyriform (γP); mesotergum divided into 3 scutal areas; posterior margin of DS convex or sub straight; scutal areas I–II unarmed (Figs. 2 – 7); area III armed with two paramedian large backwards-pointing spines (larger in some females), except in P. striatus; coxa IV with large proapical apophysis in males (Figs. 2 – 7) and females, generally larger and with curved apex in males (except P. picinguaba sp. nov., straight and equal-sized in both sexes); femur IV with (P. fuscopictus and P. striatus) or without DBA (P. bocaina sp. nov. and P. picinguaba sp. nov.); femur IV of females armed (usually with an armature different than males except P. picinguaba sp. nov.); tarsus IV with more than 16 tarsomeres; carapace dark colored and DS yellowish, with black spots or bands on scutal areas. Penis (Fig. 6) with deep cleft on VP, lateral lobes placed sub-basally; VP rounded, without globular base in lateral view; MS A + B placed forming an arch; MS B smaller than MS A (except in P. striatus); MS C with helicoidal apex; flabellum fan-like; stylus sub sigmoid, with apex inclined to the axis of stylus. Nomenclatural history. Roewer (1913) described Ampheres striatus in Caelopyginae. Later, Piza (1940) created the new genus Progonyleptoidellus, which was placed in Gonyleptinae, with a single species Progonyleptoidellus androgynus, based on a female from Alto da Serra, São Paulo. Two years later, B. Soares (1942) described the monotypic genus Laneius, for Laneius fuscopictus B. Soares, 1942, based on a female holotype and a male allotype from Boracéia, Salesópolis, São Paulo state, placing it in Gonyleptinae too. In the following year, Piza (1943) created in Gonyleptinae the new monotypic genus Piraquara, with the single species Piraquara schubarti Piza, 1943 from Serra da Piraquara, Itanhaém, São Paulo. In the same year, Roewer (1943) described Cadeadoius atroluteus Roewer, 1943 in Goniosomatinae. One year later, B. Soares (1944 a) synonymized Progonyleptoidellus and Piraquara under Gonyleptoides Roewer, 1913, a formerly monotypic genus from Rio de Janeiro. Soares & Soares (1949) synonymized Laneius with Gonazula Roewer, 1930, a formerly monotypic genus from Santa Catarina, producing the new combination Gonazula fuscopicta. In the same paper, Soares & Soares (1949: 167) synonymized Gonyleptoides with Geraeocormobius Holmberg, 1888, making the new combinations Geraeocormobius androgynus and G. schubarti; as pointed out by Acosta et al. (2007 a), the correct (original) spelling of the generic name is Geraeocormobius, while the majority of subsequent authors employed the incorrect spelling “ Geraecormobius ”. Still in the same contribution, Soares & Soares (1949) transferred Cadeadoius atroluteus to Geraeocormobius. Five years later, Soares & Soares (1954) described Geraeocormobius orguensis based on a female from Serra dos Órgãos, Teresópolis, Rio de Janeiro state. Later, Soares & Bauab-Vianna (1972) revalidated Progonyleptoidellus and considered Piraquara as its junior subjective synonym. Thirteen years later, Soares & Soares (1985) synonymized Piraquara schubarti and Cadeadoius atroluteus with Progonyleptoidellus androgynus, and revalidated Laneius. Furthermore, Soares & Soares (1985) also erected the new subfamily Progonyleptoidellinae, with Progonyleptoidellus as the type genus, and included the genera Iguapeia Mello-Leitão, 1935 b, Laneius, Heliella B. Soares, 1945 b, Cadeadoius Mello-Leitão, 1936, Moreiranula Roewer, 1930, Gonyleptoides, Opisthoplites Sørensen, 1884 and Stylopisthos Roewer, 1930. Several years later, Kury & Pinto-da-Rocha (1997) considered the monotypic genus Laneius as a junior subjective synonym of Progonyleptoidellus, resulting in a new combination, Progonyleptoidellus fuscopictus. Moreover, the authors also considered Progonyleptoidellus androgynus as a junior subjective synonym of Ampheres striatus, so they removed this species from Ampheres C. L. Koch, 1839 (Caelopyginae) and assigned it to Progonyleptoidellus (Progonyleptoidellinae), resulting in the new combination Progonyleptoidellus striatus. Five years later, Kury (2003), in his annotated catalogue, transferred Geraeocormobius orguensis to Progonyleptoidellus, without justification.
Etymology. The genus name derives from the pre-existing genus Progonyleptoides + masculine form of Latin suffix – ellus. Gender masculine. Included species. Progonyleptoidellus bocaina sp. nov., Progonyleptoidellus fuscopictus (Soares, 1942), Progonyleptoidellus picinguaba sp. nov. and Progonyleptoidellus striatus (Roewer, 1913).