Dodona formosana Matsumura 1919
- Dataset
- A distinct species, Dodonaformosana, detected in the Dodonaeugenes species complex: clarification of the taxonomic status of the Punch butterfly in Taiwan
- Rank
- SPECIES
- Published in
- Wu, Li-Wei, Lin, Wen-Jie, Hsu, Yu-Feng (2018): A distinct species, Dodonaformosana, detected in the Dodonaeugenes species complex: clarification of the taxonomic status of the Punch butterfly in Taiwan. ZooKeys 736: 59-77, DOI: http://dx.doi.org/10.3897/zookeys.736.22062, URL: http://dx.doi.org/10.3897/zookeys.736.22062
Classification
- kingdom
- Animalia
- phylum
- Arthropoda
- class
- Insecta
- order
- Lepidoptera
- family
- Riodinidae
- genus
- Dodona
- species
- Dodona formosana
diagnosis
Diagnosis. Wing pattern of Dodona formosana is similar to D. eugenes and D. maculosa Leech 1890, in sharing small markings and narrow stripes (Fig. 5), but can be distinguished by following characters: (1) The silver stripes on the underside of the hindwing is more prominent in D. eugenes (Fig. 5 E, F); the spots on the wing upper side is orange in D. formosana (Fig. 5 A-D), but light orange or yellow is presented in D. eugenes (Fig. 5 E, F). (2) The yellow spots posterior to cell M on the underside of the forewing are brighter in D. formosana (Fig. 5 A-D), the proximal band on the upper side of the hindwing is less prominent in D. formosana (Fig. 5 A-D). Male genitalia of D. formosana is quite different from that of D. eugenes and D. maculosa, especially in the following characters (Fig. 1 A-D, F): (1) the valva is narrower and longer in D. formosana, its long axis is nearly two times as long as short axis; (2) the costal process is L-shaped in D. formosana, whereas it is triangular in D. eugenes and D. maculosa; (3) the juxta of D. formosana and D. eugenes are nearly divided in the middle (Fig. 1 A-D), whereas they are strongly fused in D. maculosa (Fig. 1 F); besides, the branches of juxta are slenderer in D. formosana (Figs 1 A-B) than D. eugenes (Fig. 1 C-D); (4) the phallus is longer and slenderer in D. formosana; (5) the tip of carina penis is longer and down-curved in D. maculosa (Fig. 1 F), and bifurcated in D. eugenes (Fig. 1 C-D).
distribution
Distribution. This species is endemic in Taiwan.
materials_examined
Material examined. Holotype (Fig. 5 K). ♂ labelled ' 35.7.23 ', ' Shito [in Chinese] ' (Shito, Pinglin, Xinbei, TAIWAN), ' 972 ', ' type Dodona eugenes var. formosana Matsumura' (SEHU) Additional material. 1 ♂ labelled ' 2013. IX. 24 (Collect.) ', ' Mt. Erge Shiding Xinbei', ' leg. W. J. Lin, C. W. Huang, C. J. Peng, Y. T. Chen'; 1 ♂ labelled ' 2014. VII. 03 (Collect.) ', ' 2014. IX. 20 (Eclosion) ', ' Mt. Erge Shiding Xinbei', ' leg. W. J. Lin', ' genitalia Dn 066 '; 1 ♂ labelled ' 2014. V. 17 (Collect.) ', ' 2014. VII. 09 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. K. W. Hsiao, Y. M. Hsu', ' genitalia Dn 034 ', ' 14 E 37 '; 1 ♂ labelled ' 2014. VII. 07 (Eclosion) ', ' 2014. V. 17 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. K. W. Hsiao, Y. M. Hsu', ' 14 E 37 '; 1 ♂ labelled ' 2014. V. 17 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. K. W. Hsiao, Y. M. Hsu'; 1 ♂ labelled ' 2014. V. 17 (Collect.) ', ' 2014. VII. 09 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. K. W. Hsiao, Y. M. Hsu', ' genitalia lot. Dn 038 ', ' 14 E 37 '; 1 ♂ labelled ' 2014. V. 17 (Collect.) ', ' 2014. VII. 09 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. K. W. Hsiao, Y. M. Hsu', ' genitalia lot. Dn 039 ', ' 14 E 37 '; 1 ♂ labelled ' 2014. V. 17 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. K. W. Hsiao, Y. M. Hsu'; 1 ♂ labelled ' 2014. V. 17 (Collect.) ', ' 2014. VII. 09 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. K. W. Hsiao, Y. M. Hsu', ' genitalia lot. Dn 033 ', ' 14 E 37 '; 1 ♂ labelled ' 2014. V. 24 (Collect.) ', ' 2014. VII. 15 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin, C. P. Hseuh', ' 14 E 50 '; 1 ♂ labelled ' 2014. V. 24 (Collect.) ', ' 2014. VII. 15 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin, C. P. Hseuh', ' 14 E 50 '; 1 ♀ labelled ' 2014. VII. 09 (Eclosion) ', ' 2014. V. 24 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin, C. P. Hseuh', ' 14 E 50 '; 1 ♀ labelled ' 2014. V. 24 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin, C. P. Hseuh', ' 14 E 48 '; 1 ♀ labelled ' 2014. V. 24 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin, C. P. Hseuh', ' 14 E 48 '; 1 ♂ labelled ' 2014. V. 24 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin, C. P. Hseuh'; 1 ♂ labelled ' 2014. VIII. 03 (Collect.) ', ' Mt. Erge Shiding Xinbei', ' leg. W. J. Lin', ' genitalia lot. Dn 023 '; 1 ♂ labelled ' 2014. VIII. 03 (Collect.) ', ' 2014. IX. 25 (Eclosion) ', ' Mt. Erge Shiding Xinbei', ' leg. W. J. Lin', ' genitalia lot. Dn 024 '; 1 ♂ labelled ' 2014. VIII. 03 (Collect.) ', ' 2014. IX. 21 (Eclosion) ', ' Mt. Erge Shiding Xinbei', ' leg. W. J. Lin', ' genitalia lot. Dn 025 '; 1 ♂ labelled ' 2015. VI. 27 (Collect.) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin', ' genitalia lot. Dn 067 '; 1 ♂ labelled ' 2015. VI. 27 (Collect.) ', ' 2015. VIII. 17 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin', ' genitalia lot. Dn 064 '; 1 ♂ labelled ' 2015. VIII. 04 (Collect.) ', ' 2015. IX. 20 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin', ' genitalia lot. Dn 065 '; 1 ♂ labelled ' 2015. VIII. 04 (Collect.) ', ' 2015. IX. 24 (Eclosion) ', ' Daluntou Neihu Taipei', ' leg. W. J. Lin', ' genitalia lot. Dn 082 '; 1 ♂ labelled ' 2014. VIII. 30 (Collect.) ', ' Mt. Neinaotsui Jianshi Xinchu', ' leg. L. H. Wang, J. Y. Liang', ' genitalia lot. Dn 032 ', ' 14 H 41 - 1 - MA 03 '; 1 ♂ labelled ' 2014. VIII. 30 (Collect.) ', ' Mt. Neinaotsui Jianshi Xinchu', ' leg. L. H. Wang, J. Y. Liang', ' genitalia lot. Dn 070 ', ' 14 H 41 - 2 - MS 01 '; 1 ♂ labelled ' 2013. VII. 27 (Eclosion) ', ' 2013. VI. 20 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin'; 1 ♀ labelled ' 2013. VIII. 02 (Eclosion) ', ' 2013. VI. 22 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin'; 1 ♂ labelled ' 2013. VIII. 10 (Eclosion) ', ' 2013. VI. 25 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin', ' genitalia lot. Dn 027 '; 1 ♂ labelled ' 2013. IX. 29 (Eclosion) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin', ' genitalia lot. Dn 010 '; 1 ♂ labelled ' 2013. XI. 08 (Eclosion) ', ' 2013. XI. 29 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin', ' genitalia lot. Dn 026 '; 1 ♂ labelled ' 2014. III. 16 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin', ' genitalia lot. Dn 028 '; 1 ♂ labelled ' 2014. IV. 01 (Eclosion) ', ' 2014. III. 16 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin', ' genitalia lot. Dn 028 '; 1 ♂ labelled ' 2014. X. 07 (Eclosion) ', ' 2014. XI. 20 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin, C. C. Lin', ' genitalia lot. Dn 083 '; 1 ♂ labelled ' 2014. IV. 02 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin, C. J. Chang, Y. H. Lin, M. F. Chou', ' genitalia lot. Dn 059 '; 1 ♂ labelled ' 2014. IV. 02 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin, C. J. Chang, Y. H. Lin, M. F. Chou', ' genitalia lot. Dn 084 '; 1 ♂ labelled ' 2014. IX. 28 (Collect.) ', ' 2014. VIII. 25 (Eclosion) ', ' Songmao Forest Road Hoping', ' leg. W. J. Lin, C. W. Huang, Y. M. Hsu, Y. H. Lin', ' genitalia lot. Dn 085 '; 1 ♂ labelled ' 2014. XI. 11 (Eclosion) ', ' 2014. X. 07 (Collect.) ', ' Mt. Malun Hoping Taichung', ' leg. W. J. Lin, C. C. Lin', ' genitalia lot. Dn 068 '; 1 ♂ labelled ' 2015. VII. 05 (Collect.) ', ' Taiwan No. 8 Highway 119.5 K Xiulin Hualien', ' leg. L. Huang', ' genitalia lot. Dn 062 ', ' DNA lot. Rd 030 '.