Pupina arula arula Benson, 1856
- Dataset
- Malay Peninsular Terrestrial Molluscs
- Rank
- SPECIES
Classification
- kingdom
- Animalia
- phylum
- Mollusca
- class
- Gastropoda
- family
- Pupinidae
- genus
- Pupina
- species
- Pupina arula arula
morphology
Benson (1856) original descriptions on Pupina arula – “Testa imperforata, conoideo-ovata, longitudinaliter striatula, nitidissima, fusco-rubella, apice conoideo, acuto, sutura calloso-marginata; anfractibus 6, ultimo spiram subsequante, antice breviter ascendente; apertura circulari, angulo superiori acuto adjecto, callo parietali superne lamella intrante munito; columella profunde incisa, canalem extus apparentem, lingua lata parietali obtectum, callisque duobus divergentibus marginatum, exhibente; peristomate obtuso, expansiusculo, extus marginato, margine dextro supra medium arcuato; basi foveata. Operculo ?”
Möllendorff (1891) descriptions on syntype Pupina arula perakensis – “testa minore, graciliore, apice acutiore”
Kobelt (1902) descriptions on Pupina (Tylotoechus) arula – “Schale ungenabelt, unten grubig eingedrückt, ei-kegelförmig, gestreift, sehr glänzend, rotbraun; Apex kegelförmig, spitz; Naht schwielig, geraudet; 6 Windungen, letzte hinten so hoch wie das Gewinde, vorn leicht ansteigend; Mündung fast senkrecht, kreisrund, am oberen, scharfen Winkel erweitert, Callus oben mit einer eindringenden Lamelle, Spindelrand tief eingeschnitten, einen aussen sichtbaren Kanal bildend, der durch eine breite Zunge überdeckt und durch 2 auseinandergehende Schwielen gerandet ist; Mundrand stumpf, leicht ausgebreitet, aussen gerandet, rechts über der Mitte gebogen.”
diagnostic
“It has much affinity with P. aurea, Hinds, the superior canal being rather simulated than actually developed, and being formed by an angle, at the top of the otherwise circular aperture, cut off from the lower portion, in part, by the parietal lamina.” (Benson, 1856)
“The Perak form is smaller, 8 instead of 9 millim. In length, comparatively more slender, showing a diameter of 2; 1 instead of 9:5; the apex is more pointed.” (Möllendorff, 1891)
“A single live collected example (height 7.52 mm., diameter 5.3 mm., with 4 5/8 whorls) is referred here. Although smaller than the type of arula (height 9 mm.) from Yanglan, Tenasserim, the difference probably is not significant. Pupina lowei de Morgan, 1885 and P. siamensis (Moellendorff, 1902) (see Arch. F. Mollusk. 86, pl. 2, fig. 15) may be either subspecies or synonyms. Various sets of shells in the FMNH collection from Perak labelled lowei and arula do not seem separatable.”
distribution
Type locality – “ad Yunglaw, in valle Tenasserim” (Benson, 1856); Pupina arula perakensis “Bukit Pondong” leg. Townsend & Hungerford (Möllendorff, 1891)
Other locality – “Hill near the hot springs, about 400 m from the main road from Tandjong Rambutan to Ipoh, near Tambun, Perak” leg. Benthem-Jutting/1946 (Benthem-Jutting, 1960); “Thailand; Doi Sutep at 1,100 meters” (Solem, 1966); “limestone island C, Temenggor dam, 230 m” leg. Davison/Aug. 28, 1993 (Davison, 1995)
size
Pupina arula – “Long. 9, diam. 5 mill.” (Benson, 1856); Pupina arula perakensis – “alt. 8, diam. 4mm.” (Möllendorff, 1891); Pupina (Tylotoechus) arula – “H. 9, Durchm. 5, Mündungsweite 3 mm.” (Kobelt, 1902)
Name
- Homonyms
- Pupina arula arula Benson, 1856
Bibliographic References
- G. Nevill (1878) part I Gastropoda : Pulmonata and Prosobranchia-Neurobranchia http://borneanlandsnails.myspecies.info/node/85